Gregarious in coniferous woods, mainly under
Picea. Autumn. Occasional, but locally common in the Nordic countries. Also recorded in Germany and Scotland. Widely distributed in Norway, but not very common. Most records in the south-east part of the country.
Pileus 5-22 mm across,
conical, campanulate to convex, sometimes with an umbo,
sulcate, translucent-striate, at first white-pruinose and
then often looking almost bluish black, glabrescent, dark
sepia brown at the centre, paler to whitish towards the
margin. Lamellae 17-20 reaching the stipe,
ascending, narrowly adnate with a decurrent tooth, pale
grey or grey with a brownish shade, the edge paler. Stipe
25-80 x 0.5-2 mm, hollow, terete, straight to somewhat curved,
equal, pruinose above, glabrous below, whitish to greyish
above, pale brown below. Odour of Pelargonium
graveolens, sometimes indistinctive.
Basidia 25-40 x 6-8, slender-clavate, 4-spored, with sterigmata 3-5 μm long. Spores (7.5-)8-9(-10.5) x (3.5-)4-5.1 μm, Q 1.6-2.1, Qav≈1.9, pip shaped, amyloid. Cheilocystidia
12-46 x 7-18 μm, forming a sterile band, saccate or more or less shaped like
an hour-glass, with broad base, some more clavate, obpyriform
or spheropedunculate, covered with few to more numerous
warts or cylindrical, branched and curved, fairly coarse excrescences,
1-10 x 1-2 μm. Pleurocystidia similar. Hyphae of the pileipellis 1.5-3.5 μm wide, covered with straight to curved or much branched excrescences 1-20 x 1-2 μm,
forming dense, corralloid masses. Hyphae
of the cortical layer of the stipe 1.5-3.5 μm wide, smooth to diverticulate, excrescences 0.5-4 x 1-1.5 μm, the terminal cells up to 10 μm wide, variously inflated, covered with fairly coarse excrescences. Clamp connections present at all tissues.
Microphotos of cheilocystidia.
Mycena septentrionalis was a well
known species in Scandinavia, known as Mycena sepia
sensu Lundell. Maas geesteranus (1984), however, showed
that the original M. sepia was another species
and proposed the new name M. septentrionalis. He
stressed the resemblance between M. septentrionalis
and M. filopes and focused
on several features to separate the two species: In M.
septentrionalis the cheilocystidia are mostly covered
with unevenly spaced, coarse, often curved, simple to branched
excrescences, and many cystidia are saccate or shaped like
an hour-glass, with the apical part and the base equally
broad. In M. filopes
the cheilocystidia are mostly covered with evenly spaced,
cylindrical, narrow excrescences, not shaped like an hour-glass.
The terminal cells of the stipe cortex can also be used
to separate the two species. In M. septentrionalis
they are covered with fairly coarse excrescences, while
they are covered with fine and slender ones in M. filopes.
In addition he mentioned the odour, which is of iodoform
when drying out in M. filopes
and pungent-aromatic, of Pelargonium graveolens
in M. septentrionalis.
In my opinion Maas Geesteranus exaggerated
the problem. There are small chances to mix up the two species. Probably, M. metata is more likely to be confused with M. septentrionalis. They often grow together, but that shoudn't be a problem either. M. septentrionalis should be readily recognized
in the field on account of the dark, sepia brown pileus,
the grey to brownish lamellae with a decurrent tooth, and
the smell of Pelargonium. The odour may be absent,
but according to my experience it is quite constant. Microscopically
the hour-glass shaped cheilocystidia is a reliable character.
An equally dark brown species is M. rapiolens, but this has large, clavate cheilocystidia that are not constricted, much wider hyphae of the pileipellis and very rare or absent clamps.
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